Faculty Directory

EDUCATION

PhD University of Pennsylvania, 1960

I became an emeritus professor in 2005 and closed my laboratory. I therefore am no longer actively involved in research nor am I accepting new graduate students. I do, however, remain active in the teaching programs of the Psychology Department and the Neuroscience and Cognitive Science program.

• Study of motivation in animals
• Vision of birds and how visual information is processed in the avian central nervous system
• Determination of the central visual pathways in the nervous systems of birds
• Application of various behavioral psychophysical techniques to measure (in pigeons, quail, hawks, owls, starlings, and other birds) :
  • Visual acuity
  • Luminance differences
  • Spatial and temporal contrast sensitivity
• Physiological optics of the eyes of various birds
• Evolution of the brain 
• Evolution of behavior, comparative neuroanatomy, and animal intelligence